Research Positions

2003- Associate Professor (tenured), Coastal Oregon Marine Experiment Station and Department of Fisheries and Wildlife, Oregon State University.

2003- Scientific Adviser, Norwegian Institute for Nature Research (NINA), Tungasletta 2, N-7485 Trondheim, Norway.

2001-02Associate Professor (tenure track), Coastal Oregon Marine Experiment Station and Department of Fisheries and Wildlife, Oregon State University.

2001-02 Adjunct Professor, Norwegian Institute for Nature Research (NINA), Tungasletta 2, N-7485 Trondheim, Norway.

1999-00. Research Scientist (I), Division for Aquatic Ecology, Norwegian Institute for Nature Research.

1999-02. Adjunct Professor, Department of Biology, University of Waterloo, Waterloo, Canada.

1998. Guest Researcher. Institute of Ecology, Department of Zoology, University of Göteborg, Göteborg (Gothenburg), Sweden.

1994-98 Research Scientist (II), Norwegian Institute for Nature Research, Trondheim, Norway.

1991-93 Postdoctoral Visiting Scientist, Norwegian Institute for Nature Research, Trondheim, Norway.

Degrees

Ph.D. Department of Zoology, University of Toronto, Toronto, Canada. 1991
M.Sc. Department of Biological Sciences, Simon Fraser University, Burnaby, Canada. 1986
B.Sc. Department of Biology, Queen's University, Kingston, Canada. 1983

Research Interests

Behavioural and evolutionary ecology of fishes, with an emphasis on breeding system evolution, life history diversity, maternal effects, survival strategies and habitat, cultured and wild fish interactions, fisheries, and conservation. Interest in marine and freshwater fishes, with focus on salmonids and groundfish.

Refereed Publications

Einum, S., Fleming, I.A., Côté, I.M. and Reynolds, J.R. Population stability in salmon species: Effects of population size and female reproductive allocation. Journal of Animal Ecology, in press.

Fleming, I.A. and Reynolds, J.D. Salmon breeding systems. In A.P. Hendry and S.C. Stearns. Evolution in Salmon. Oxford University Press, Oxford, USA, in press.

Hindar, K. and Fleming, I.A. Behavioural and genetic interactions between escaped farm and wild Atlantic salmon. In T.M. Bert and J.L. Nielsen (eds.) Ecological and genetic implications of aquaculture activities. Kluwer Academic, New York, in press.

Garant, D.*, Fleming, I.A.*, Einum, S.* and Bernatchez, L. 2003. Alternative male life-history tactics as potential vehicles for speeding introgression of farm salmon traits into wild populations. Ecology Letters 6: 541-549. (* contributed equally to the work)

Forseth, T., Ugedal, O., Jonsson, B. and Fleming, I.A. 2003. Ecological character displacement in Arctic charr caused by competition from brown trout. Oikos 101: 467-478.

Einum, S. and Fleming, I.A. 2002. Does within-population variation in fish egg size reflect maternal influences on optimal values? American Naturalist 160: 756-765.

Lembo, G., Spedicato, M.T., Økland, F., Carbonara, P., Fleming, I.A., McKinley, R.S., Thorstad, E.B., Sisak, M. and Ragonese, S. 2002. A wireless communication system for determining site fidelity of juvenile dusky groupers, Epinephelus marginatus (Lowe, 1834), using coded acoustic transmitters. Hydrobiologia 483: 249-257.

Einum, S., Hendry, A.P. and Fleming, I.A. 2002. Egg size evolution in aquatic environments: does oxygen availability constrain size? Proceeding of the Royal Society, London B 269: 2325-2330.

Fleming, I.A., Agustsson, T., Finstad, B., Johnsson, J. and Björnsson, B.Th. 2002. Effects of domestication on growth physiology and endocrinology of Atlantic salmon. Canadian Journal of Fisheries and Aquatic Sciences 59: 1323-1330.

Singer, T.D., Clements, K.M., Semple, J., Schulte, P.M., Bystriansky, J., Finstad, B., Fleming, I.A. and McKinley, R.S. 2002. Seawater tolerance and gene expression in Atlantic salmon smolts: a comparison of hatchery-reared farm and native strains. Canadian Journal of Fisheries and Aquatic Sciences 59: 125-135.

Armstrong, J.D., Einum, S., Fleming, I.A. and Rycroft, P. 2001. A method for tracking the behaviour of mature and immature parr around nests during spawning. Journal of Fish Biology 59: 1023-1032.

Fleming, I.A. and Petersson, E. 2001. The ability of released, hatchery salmonids to breed and contribute to the natural productivity of wild populations. Nordic Journal of Freshwater Research 75: 71-98.

Einum, S. and Fleming, I.A. 2001. Implications of stocking: ecological interactions between wild and released salmonids. Nordic Journal of Freshwater Research 75: 56-70.

Johnsson, J.I., Höjesjö, J. and Fleming, I.A. 2001. Behavioural and heart rate responses to predation risk in wild and domesticated Atlantic salmon. Canadian Journal of Fisheries and Aquatic Sciences 58: 788-794.

Fleming, I.A., Hindar, K., Mjølnerød, I.B., Jonsson, B., Balstad, T. and Lamberg, A. 2000. Lifetime success and interactions of farm salmon invading a native population. Proceedings of the Royal Society, London B 267: 1517-1524.

Einum, S. and Fleming, I.A. 2000. Highly fecund mothers sacrifice offspring survival to maximise fitness. Nature 405: 565-567.

Einum, S. and Fleming, I.A. 2000. Selection against late emergence and small offspring in Atlantic salmon. Evolution 54: 628-639.

Lembo, G., Fleming, I.A., Økland, F., Carbonara, P. and Spedicato, M.T. 1999. Site fidelity of the dusky grouper Epinephelus marginatus (Lowe, 1834) studied by hydroacoustic telemetry. Marine Life 9(2): 37-43.

Einum, S. and Fleming, I.A. 1999. Maternal effects of egg size in brown trout (Salmo trutta): norms of reaction to environmental quality. Proceedings of the Royal Society, London B 266: 1995-2000.

Lembo, G., Fleming, I.A., Økland, F., Carbonara, P. and Spedicato, M.T. 1999.Homing behaviour and site fidelity of Epinephelus marginatus around the island of Ustica: preliminary results from a telemetry study. Biologia Marina Mediterranea 6: 90-99.

Fleming, I.A. 1998. Pattern and variability in the breeding system of Atlantic salmon, with comparisons to other salmonids. Canadian Journal of Fisheries and Aquatic Sciences 55(Suppl. 1): 59-76.

Armstrong, J.D., Grant, J.W.A., Forsgren, H.L., Fausch, K.D., DeGraaf, R.M., Fleming, I.A., Prowse, T.D. and Schlosser, I.J. 1998. The application of science to the management of Atlantic salmon: integration across scales. Canadian Journal of Fisheries and Aquatic Sciences 55(Suppl. 1): 303-311.

Lacroix, G.L. and Fleming, I.A. 1998. Ecological and behavioural interactions between farmed and wild Atlantic salmon: consequences for wild salmon. Canadian Stock Assessment Secretariat. Research Document 98/162.

Lembo, G., Fleming, I.A., Økland, F., Carbonara, P., Ragonese, S. and Spedicato, M.T. 1998. Design of a telemetry study for the dusky grouper (Epinephelus marginatus, Lowe 1834). Biologia Marina Mediterranea 5: 1258-1265.

Mjølnerød, I.B., Fleming, I.A., Refseth, U.H. and Hindar, K. 1998. Mate and sperm competition during multiple-male spawnings of Atlantic salmon. Canadian Journal of Zoology 76: 70-75.

Fleming, I.A., Lamberg, A. and Jonsson, B. 1997. Effects of early experience on the reproductive performance of Atlantic salmon. Behavioral Ecology 8: 470-480.

Fleming, I.A. and Einum, S. 1997. Experimental tests of genetic divergence of farmed from wild Atlantic salmon due to domestication. ICES Journal of Marine Science 54: 1051-1063.

Einum, S. and Fleming, I.A. 1997. Genetic divergence and interactions in the wild among native, farmed and hybrid Atlantic salmon. Journal of Fish Biology 50: 634-651.

Fleming, I.A. 1996. Reproductive strategies of Atlantic salmon: ecology and evolution. Reviews in Fish Biology and Fisheries 6: 379-416.

Fleming, I.A., Jonsson, B., Gross, M.R. and Lamberg, A. 1996. An experimental study of the reproductive behaviour and success of farmed and wild Atlantic salmon (Salmo salar). Journal of Applied Ecology 33: 893-905.

Jonsson, N., Jonsson, B. and Fleming, I.A. 1996. Phenotypically plastic response of egg production to early growth in Atlantic salmon, Salmo salar. Functional Ecology 10: 89-96.

Fleming, I.A. 1994. Captive breeding and the conservation of wild salmon populations. Conservation Biology 8: 886-888.

Fleming, I.A., and Gross, M.R. 1994. Breeding competition in a Pacific salmon (coho: Oncorhynchus kisutch): measures of natural and sexual selection. Evolution 48: 637-657.
[This paper received the W.F. Thompson Award, American Institute of Fishery Research Biologists.]

Fleming, I.A., Jonsson, B. and Gross, M.R. 1994. Phenotypic divergence of sea-ranched, farmed and wild salmon. Canadian Journal of Fisheries and Aquatic Sciences 51: 2808-2824.

Fleming, I.A. and Gross, M.R. 1993. Breeding success of hatchery and wild coho salmon (Oncorhynchus kisutch) in competition. Ecological Applications 3: 230-245.

Fleming, I.A. and Gross, M.R. 1992. Reproductive behaviour of hatchery and wild coho salmon (Oncorhynchus kisutch): does it differ? Aquaculture 103: 101-121.

Fleming, I.A. and Gross, M.R. 1990. Latitudinal clines: a tradeoff between egg number and size in Pacific salmon. Ecology 71: 1-11.

Fleming, I.A. and Gross, M.R. 1989. Evolution of adult female life history and morphology in a Pacific salmon (Coho: Oncorhynchus kisutch). Evolution 43: 141-157.

Fleming, I.A. and Ng, S. 1987. Evaluation of techniques for fixing, preserving and measuring salmon eggs. Canadian Journal of Fisheries and Aquatic Sciences 44: 1957-1962.

Fleming, I.A. and Johansen, P.H. 1984. Density and aggressive behaviour in young-of-the-year largemouth bass (Micropterus salmoides). Canadian Journal of Zoology 62: 1454-1455.

Book Chapters

Fleming, I.A. and Jensen, A.J. 2001. Fisheries: Effects of climate change on the life cycles of salmon, pp. 309-312. In I. Douglas (ed.) Causes and consequences of global environmental change. Volume 3. Encyclopedia of Global Environmental Change. Wiley, Chichester. (ISBN 0-471-97796-9)

Fleming, R.A., Fleming, R.L. and Fleming, I.A. 2001. Life cycles, pp. 385-389. In H.A. Mooney and J. Canadell (eds.) The Earth System: biological and ecological dimensions of global environmental change. Volume 2. Encyclopedia of Global Environmental Change. Wiley, Chichester. 625 p. (ISBN 0-471-97796-9)

Økland, F., Fleming, I.A., Thorstad, E.B., Finstad, B., Einum, S. and McKinley, R.S. 2000. EMG telemetry to record the intensity of swimming- and breeding-related behaviour in Atlantic salmon, pp. 51-58. In Moore, A. and Russell, I.A. (eds.) Advances in Fish Telemetry. Centre for Environment, Fisheries and Aquaculture Science, Lowestoft, U.K., 264 p.

Fleming, I.A. 1998. Spawning of Atlantic salmon, pp. 127-141. In R.V. Kazakov (ed.) Atlantic salmon. Nauka, St. Petersburg, Russia.575 p. (ISBN 5-02-026113-0) (in Russian)

McNeely, J.A. et al. (Contributor). 1995. Human influences on biodiversity, pp. 711-821. In V.H. Heywood (ed.) Global Biodiversity Assessment. United Nations Environmental Programme and Cambridge University Press, Cambridge. (ISBN 0-521-56481-6)

Fleming, I.A. 1995. Reproductive success and the genetic threat of cultured fish to wild populations, pp. 117-135. In D.P. Philipp, J.M. Epifanio, J.E. Marsden and J.E. Claussen (eds.). Protection of aquatic biodiversity. Proceedings of the World Fisheries Congress, Theme 3. Oxford and IBH Publishing, New Delhi, India. (ISBN 81-204-0971-X)

Jonsson, B., and Fleming, I.A. 1993. Enhancement of wild salmon populations, pp. 209-242. In G. Sundnes (ed.) Human Impact on Self-recruiting Populations. An International Symposium of the Royal Norwegian Society of Sciences and Letters, Kongsvoll, Norway, 7-11 June 1993. Tapir Press, Trondheim, Norway. (ISBN 82-519-1455-8)

Books

National Research Council (co-author). 2002. Genetic Status of Atlantic Salmon in Maine. Interim Report from the Committee on Atlantic Salmon in Maine. National Academy Press, Washington, DC. 62 pp. (ISBN 0-309-08311-7)

Thorstad, E., Fleming, I.A. and Næsje, T. (eds.) 2002. Aquatic Telemetry. Proceedings of the Fourth Conference on Fish Telemetry in Europe. Kluwer Academic. (ISBN1-4020-0877-5) (reprinted from Hydrobiologia volume 483; see below)

Edited Volumes

Thorstad, E., Fleming, I.A. and Næsje, T. (eds.) 2002. Proceedings of the Fourth Conference on Fish Telemetry in Europe. Hydrobiologia 483: 1-292. (ISSN 0018-8158)

Fleming, I.A. (ed.) 2001. Proceedings of the Workshop on the Release of Salmonid Fishes in Norway. Nordic Journal of Freshwater Research 75(1): 1-152. (ISSN 1100-4096)

Other Miscellaneous Publications

Finstad, B., McKinley, R.S., Fleming, I.A., Bjørnsson, B.Th., Augustsson, T. and Johnsson, J.I. 2002. Comparisons of wild and cultured Atlantic salmon: a case study from Norway. In McKinley, R.S., Driedzic, W.R. and MacKinlay, D. (eds.) Behavioral and Physiological Comparisons of Cultured and Wild Fish. Proceedings of the International Congress on the Biology of Fish, University of British Columbia, Canada, July 21-26, 2002. American Fisheries Society.

Fleming, I.A. 2002. Can hatchery fish contribute to the natural productivity of wild populations? pp. 32-56. In C. Orr, P. Gallaugher and J. Penikett (eds.) Speaking for the Salmon. Workshop Proceedings on Hatcheries and the Protection of Wild Salmon. Continuing Studies in Science, Simon Fraser University, Burnaby, Canada. (ISBN 0-86491-230-7)

Forseth, T., Ugedal, O., Jonsson, B., Fleming, I.A. and Gåsdal, O. 2002. How do we get large fish? pp. 27-31. In H.C. Pedersen and B. Jonsson (eds.) NINAs Strategic Institute Programmes 1996-2000: Sustainable exploitation of populations. Final Report. NINA Temahefte 18, 92 p. (ISBN 82-426-1285-4) (In Norwegian with an English abstract).

Strand, R., Fleming, I.A. and Johnsen, B.O. 2001. Releases of salmonid fishes. Kongsvoll Workshop 2000. NINA Fagrapport 045: 1-49. (ISBN 82-426-1185-8) (In Norwegian with an English abstract)

Fleming, I.A. 2001. Captive breeding: conflict between conserving genetic variation and natural behaviour, pp. 34-35. In K. Hindar and B. Jonsson (eds.) NINAs Strategic Institute Programmes 1996-2000: Conservation of biodiversity - from genes to landscapes. Final Report. NINA Temahefte 17, 125 p. (ISBN 82-426-1176-9) (In Norwegian with an English abstract).

Hindar, K., Fleming, I.A., Hansen, L.P. and Balstad, T. 2001. Hybridization between Atlantic salmon and brown trout, pp. 43-44. In K. Hindar and B. Jonsson (eds.) NINAs Strategic Institute Programmes 1996-2000: Conservation of biodiversity - from genes to landscapes. Final Report. NINA Temahefte 17, 125 p. (ISBN 82-426-1176-9) (In Norwegian with an English abstract).

Fleming, I.A., Hindar, K., Jonsson, B. and Hansen, L.P. 2001. Effects of escape salmon on wild salmon, pp. 94-97. In K. Hindar and B. Jonsson (eds.) NINAs Strategic Institute Programmes 1996-2000: Conservation of biodiversity - from genes to landscapes. Final Report. NINA Temahefte 17, 125 p. (ISBN 82-426-1176-9) (In Norwegian with an English abstract).

Berger, H.M., Lamberg, A., Fleming, I.A., Hindar, K. and Fjeldstad, H.-P. 2001. Reconstruction of spawning sites for anadromous brown trout (Salmo trutta L.) and Atlantic salmon (Salmo salar) in the river Gråelva, Stjørdalselva watershed in central Norway during 1999-2000. NINA Oppradsmelding 678, 27 p. (ISBN 82-426-1202-1) (In Norwegian with an English abstract).

Fleming, I.A. and Einum, S. 2000. The implications of maternal decisions for offspring growth and survival, pp. 63-66. In Letcher, B. and MacKinlay, D. (eds.) Consequences of Salmonid Growth Variability. Proceedings of the International Congress on the Biology of Fish, University of Aberdeen, Scotland, July 23-27, 2000. American Fisheries Society. (ISBN 1-894337-16-6).

Fleming, I.A. 2000. What determines the ability of farm salmon to invade wild populations? pp. 23-24. Gallaugher, P. and C. Orr (eds.) Speaking for the Salmon. Workshop Proceedings on Aquaculture and the Protection of Wild Salmon. Continuing Studies in Science, Simon Fraser University, Burnaby, Canada. (ISBN 0-86491-218-8)

Fleming, I.A. 1999. Behavioural studies in salmonid fishes. In M.M. Hansen (ed.) Proceedings of the Second Workshop of the European Concerted Action on Identification , Management and Exploitation of Genetic Resources in Brown Trout (Salmo trutta), St. Feliu de Guixols, Spain, 28 June - 4 July 1999.

Lund, R.A., Finstad, B., Fleming, I.A. and Hindar, K. 1999. Oppdrettslaks og ville bestander av laksefisk. (Farmed salmon and wild populations of salmonid fishes), pp. 133-147. In K.H. Brox (ed.) Brennpunkt Natur: Aktuelt om natur og miljø. (Focal Point Nature: news about nature and environment) Tapir, Trondheim (in Norwegian) ISBN 82-519-1551-1.

Fleming, I.A. 1997. Evolution and the Aquatic Ecosystem: Defining Unique Units in Population Conservation. J.L. Nielsen (ed.). Fisheries Management and Ecology 4: 253-254.

Forseth, T., Halvorsen, G.A., Ugedal, O., Fleming, I.A., Schartau, A.K.L., Nøst, T., Hartvigsen, R., Raddum, G., Mooji, W. and Kleiven, E. 1997. Biological status of limed lakes. NINA Oppdragsmelding 508: 1-52. (In Norwegian with an English abstract)

Forseth, T., Halvorsen, G.A., Ugedal, O., Fleming, I.A., Schartau, A.K.L., Nøst, T., Hartvigsen, R., Raddum, G., Mooji, W. and Kleiven, E. 1997. Biological status of limed lakes - report for the individual lakes. NINA Oppdragsmelding 509: 1-232.

Økland, F., Lamberg, A., Heggberget, T.G., Fleming, I.A. and McKinley, R.S. 1996. Identification of spawning behaviour in Atlantic salmon (Salmo salar L.) by radiotelemetry, pp. 35-46. In E. Baras and J.C. Philippart (eds.) Underwater Biotelemetry, Proceedings of the First Conference and Workshop on Fish Telemetry in Europe, University of Liège, Belgium.

Hindar, K., Fleming, I.A., Jonsson, N., Breistein, J., Sægrov, H., Karlsbakk, E., Gammelsæter, M. and Dønnum, B.O. 1996. Rainbow trout in Norway: occurrence, reproduction and establishment. NINA Research Report 454: 1-32. (In Norwegian with an English abstract)

Jonsson, N., Jonsson, B. and Fleming, I.A. 1996. Growth rate of juvenile salmon influences their egg production as adults. Naturen 120: 255-258. (In Norwegian)

Fleming, I.A. and Aagaard, K. 1993. Documentation and measurement of biodiversity. Report to the United Nations Environmental Program Expert Conference on Biodiversity, Norway, May 1993. NINA Utredning 50: 1-23.

Jonsson, B., Andersen, R., Hansen, L.P, Fleming, I.A. and Bjørge, A. 1993. Sustainable use of biodiversity. Report to the United Nations Environmental Program Expert Conference on Biodiversity, Norway, May. NINA Utredning 48: 1-22.

Fleming, I.A. and Hindar, K. 1992. When hot geology meets cold climate on an island. Trends in Ecology and Evolution 7: 360-361.

Fleming, I.A. 1990. Breeding competition and reproductive success: hatchery versus wild coho salmon, pp. 63-70. In: Changing perspectives: stocking as a management tool in Ontario. Proceedings of the Second Annual Professional Workshop, American Fisheries Society, Southern Ontario Chapter.

Awards and Fellowships

2002. Outstanding Service Award, Department of Fisheries and Wildlife, Oregon State University.
1998. Mobility Fellowship for Senior, Established Researcher, Nordic Academy for Advanced Study.
1996. W.F. Thompson Award, American Institute of Fishery Research Biologists.
(Outstanding student publication in fisheries for 1994 [see Fleming and Gross 1994 above].)
1984-91. Graduate Fellowships: Natural Sciences and Engineering Research Council of Canada Postgraduate Scholarship, University of Toronto Open Fellowship, Ontario Graduate Scholarship, British Columbia Special Graduate Research Fellowship, British Columbia Post-Secondary Scholarship, Canadian Wildlife Foundation Scholarship, Fishery Scholarship - British Columbia Packers Limited, Theodore Roosevelt Memorial Fund - American Museum of Natural History.

Research Grants and Contracts

2003-06. "Evaluation of emergent structure in low-relief benthic habitats as a criterion for defining the essential habitat of juvenile Alaskan flatfishes." North Pacific Research Board. (PI )
2003-06. "Quantifying biological risks of growth-enhanced transgenic salmon." Norwegian Research Council. (Associate PI)
"Evolutionary and cascading ecological effects of translocation of native species into new ecosystems." Norwegian Research Council. (Associate PI)
2002-03. "A pilot study for evaluating individual behavior, residency, and growth of juvenile chinook salmon in estuarine marsh-channel networks." Oregon Sea Grant, Program Development. (PI)
2003-04. "Potential impacts of molluscan shellfish culture on salmon in estuaries." Western Regional Aquaculture Center (USDA). (PI)
2002-03. "Defining micro-habitat for juvenile rockfish off the Oregon coast: the role of abiotic and biotic structure." Mamie Markham Research Awards. (PI)
2002-03. "Genetic and ecological inference on ranges and limits of stock structure among four rockfish life-history types." Cooperative Institute for Marine Resource Studies. (co-PI)
2003-06. "Salmonid use of restored estuarine wetlands: regional application of the Salmon River Estuary Study." Oregon Sea Grant. (co-PI)
2001-04. "Impact of aquaculture on the immune response genes of natural salmonid populations: spatial and temporal genetic signatures and potential fitness consequences." European Commission Shared-Cost Research Grant. (co-PI)
2001-03. "Quantifying long-term ecological and genetic effects of interactions between farm and wild salmon." Norwegian Research Council (PI)
2000-03. "Genetic and ecological interactions between farm and wild Atlantic salmon." Canadian Network in Aquaculture, Networks of Centres of Excellence, Canada (co-PI)
2000 "Workshop on hatchery releases of anadromous salmonids." Norwegian Research Council; Directorate for Nature Management; Federation of Hydropower Companies. (PI)
2000-02. "A coordinated approach towards development of a scientific basis for management of wild Atlantic salmon in the North-east Atlantic." European Commission Concerted Action Grant. (co-PI)
1998-01. "Performance and ecological impacts of introduced and escaped fish: physiological and behavioural mechanisms." European Commission Shared-Cost Research Grant. (PI)
1998-01. "Captive breeding: conflict between maintaining genetic variation and behavioural fitness." Norwegian Research Council. (co-PI)
1998-99. "Egg production in fishes: implications for fisheries management and conservation." The British Council and The Research Council of Norway. (PI)
1997-01. "Evolution of egg size and breeding time in relation to the life history of fishes, with special reference to Atlantic salmon." Norwegian Research Council. (co-PI)
1997-9. "Ecological and genetic consequences of escaped farmed salmon for wild populations." Norwegian Research Council. (PI)
1996. "Interbreeding between escaped farmed and wild Atlantic salmon." Norwegian Directorate for Nature Management. (PI)
1995-6. "Rainbow trout in Norway: occurrence, reproduction and establishment." Norwegian Directorate for Nature Management. (co-PI)
1994-6. Hybridisation between escaped farmed Atlantic salmon and brown trout: frequency, distribution, behavioural mechanisms and effects on fitness. Commission of The European Communities Research Grant. (co-PI)
1994. "Artificial propagation: a threat to intraspecific biodiversity and a constraint on ex situ conservation." Norwegian General Scientific Research Council. (PI)
1993-4. "Understanding the threat of cultured salmon to wild populations: competition, reproductive success, and introgression." Norwegian Fisheries Research Council. (PI)
1989-90. "Breeding competition and reproductive success in hatchery versus wild coho salmon. II The role of density." Department of Fisheries and Oceans Canada, and National Science and Engineering Research Council of Canada Subvention Grant. (co-PI)
1989. "Morphological, life history and genetic response to the Great Lakes: coho salmon in the Credit River, Lake Ontario." Ontario Federation of Anglers and Hunters Sport Fisheries Research Grant. (PI)

Teaching Experience

2003. Special Topics, Estuarine and Ocean Salmon Ecology (FW 599)
2002. Ecology and Management of Marine Fishes (FW 420/520)
2000. Examiner: Graduate Courses in (1) Behavioural Ecology and Conservation and (2) Evolutionary Ecology, Norwegian University of Science and Technology.
1999. Graduate Seminar in Evolutionary Genetics, Norwegian University of Science and Technology.
1986-01. Various guest lectures in advanced courses in evolutionary theory, behavioural ecology, population dynamics, fisheries biology and conservation biology at Simon Fraser University, University of Toronto, Norwegian University of Science and Technology and Oregon State University.

Graduate Student Supervision

Ph.D.
2003- Geoff Hosack, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA.
1998- Åslaug Viken (part time), Zoological Institute, Norwegian University of Science and Technology, Trondheim, Norway. (co-supervision)
1997-00. Sigurd Einum, Zoological Institute, Norwegian University of Science and Technology, Trondheim, Norway.

M.Sc.
2003- Jena Lemke, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA. (co-supervision)
2003- Megan Petrie, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA. (co-supervision)
2002- Lance Campbell, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA. (co-supervision)
2002- David Hering, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA. (co-supervision)
2002- Julie Henning, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA. (co-supervision)
2002- Lisa Krentz, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA. (co-supervision)
2002- Marion Mann, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA.
2001- Jaclyn Richards, Department of Fisheries and Wildlife, Oregon State University, Corvallis, USA.
2001- Laura Weir, Department of Biology, University of Dalhousie, Halifax, Canada. (co-supervison)
1997-9. Sean Gillard, Zoological Institute, Norwegian University of Science and Technology, Trondheim, Norway.
1995-6. Sigurd Einum, Zoological Institute, Norwegian University of Science and Technology, Trondheim, Norway.

Visiting Researchers in the Laboratory

2002-3. Yusuke Koseki, Postdoctoral Scientist, Hokkaido University, Japan (1 year)
2001. Sigurd Einum, Visiting Scientist, Norwegian Institute for Nature Research, Trondheim, Norway (3 months).
2000. Dany Garant, Ph.D Candidate, University of Laval, Canada (4 months).

Invited Presentations

2003. Ocean Sciences Centre, Memorial University of Newfoundland, St. John's, Canada; Centre for Ecology, Evolution and Conservation, University of East Anglia, Norwich, UK.
2002. National Marine Fisheries Service, Seattle, USA; International Workshop on Salmon Farming in Chile: Keys to Sustainable Management. Puerto Montt, Chile; Taking stock in 2002: the state of the science on hatchery and wild fish issues. Annual Meeting of the North Pacific International Chapter, American Fisheries Society. Spokane, WA, USA; Hatcheries: What have we learned from out past practices? Where do we go from here? Annual Meeting of the Oregon Chapter, American Fisheries Society, Sun River, OR, USA.
2001. Department of Zoology, Oregon State University, Corvallis, OR, USA; School of Aquatic and Fisheries Sciences, University of Washington, Seattle, WA, USA; Workshop on Hatcheries & the Protection of Wild Salmon. Simon Fraser University, Vancouver, Canada.
2000. Department of Animal Ecology, Evolutionary Biology Centre, University of Uppsala, Uppsala, Sweden; Symposium on the Future of Wild Salmon and Aquaculture in Iceland. Sponsored by the Institute of Freshwater Fisheries, Reykjavik, Iceland; Brattøra Research Center, Norwegian University of Science and Technology, Trondheim, Norway; Department of Biology, University of Aarhus, Aarhus, Denmark; Department of Fisheries and Wildlife, Oregon State University, USA; Hatfield Marine Science Center, Newport, OR, USA; Workshop on Aquaculture and the Protection of Wild Salmon. Simon Fraser University, Vancouver, Canada.
1999 Department of Biology, University of Lecee, Italy; School of Biological Sciences, University of East Anglia, Norwich, England; EU workshop on identification, management and exploitation of genetic resources in brown trout. Girona, Spain; Special workshop on the Interaction between wild and farmed Atlantic salmon in the Maritime Provinces. Canadian Department of Fisheries and Oceans, Moncton, Canada;
1998. Department of Zoology, University of Göteborg, Göteborg, Sweden; Department of Zoology, University of Guelph, Guelph, Canada; Department of Zoology, Norwegian University of Science and Technology, Trondheim, Norway.
1997. Department of Biology, University of New Brunswick, Fredericton, Canada; International workshop on Integrating across scales: predicting patterns of change in Atlantic salmon. Sponsored by National Science Foundation, USA.
1995. Department of Zoology, University of Trondheim, Norway; Department of Biology, McGill University, Montreal, Canada.
1994. Department of Zoology, University of Stockholm, Sweden; Institute for Freshwater Research, Drottningholm, Sweden; Department of Biology, Dalhousie University, Halifax, Canada.
1993. Colloquium on Conservation Biology of Endangered Pacific Salmonids: Life History, Genetics and Demography. Bodega Marine Laboratory, University of California, USA; Department of Integrative Biology, University of California, Berkeley, USA; University of Northern British Columbia, Prince George, Canada; Department of Animal Ecology, University of Bergen, Bergen, Norway; Marine Fisheries Institute, Bergen, Norway.
1989-92. Department of Aquaculture, Swedish University of Agricultural Sciences, Umeå, Sweden; Zoological Institute, University of Trondheim, Trondheim, Norway; Symposium on Biological Issues in the Protection of Aquatic Biodiversity. World Fisheries Congress, Athens, Greece; Department of Biology, York University, Toronto, Canada; American Fisheries Society, Southern Ontario Chapter Annual Workshop, Dorset, Canada; Symposium on Genetics and Management: A Joint Imperative for the Future. American Fisheries Society Annual Meeting, Anchorage, USA.

Professional Activities

2003. External Examiner, Ph.D. dissertation, Umeå University, Sweden.
2003. Co-organizer, Symposium on Maternal Effects in Fish. Early Life History Section of the American Fisheries Society, Santa Cruz, August 2003.
2003. Co-organizer, Symposium on Interactions of Hatchery and Wild Fishes in Marine and Estuarine Environments. Western Division American Fisheries Society, San Diego, 14-17 April 2003.
2002-03. Steering Committee, Quantitative/Experimental Genetics Workshop, European Union funded SALGEN project.
2003. Scientific Committee, Fifth Conference on Fish Telemetry held in Europe, Ustica, Italy, June 9-13.
2003. Steering Committee, American Fisheries Society Symposium and Workshop entitled "Propagated aquatic organisms in and for aquatic resource management."
2002-03. Co-chair, Scientific Advisory Board, Cooperative Institute for Marine Resources Studies, Oregon State University.
2002. Invited Expert, Workshop on How do Aquaculture and Capture Fisheries Interact? The Ecological, Economic and Policy Implications of Aquaculture Escapes, Communication Partnership for Science and the Sea, Stanford University, California.
2002. External Examiner, Ph.D. dissertation, Aarhus University, Denmark..
2002. Invited Participant, Workshop on Measurable Salmonid Recovery Criteria, Corvallis, Oregon.
2002. Invited Expert, Information Systems for Biotechnology (ISB) workshop on Environmental Risk Assessment Modeling, Virginia Beach, Virginia.
2001-03. Member, National Academy of Sciences Review Panel for the Status of Atlantic Salmon Populations in Maine.
2001-03. Independent Science Community Member on the Northwest Power Planning Council Artificial Production Advisory Committee, Pacific Northwest, USA.
2001-02. Member, Scientific Advisory Board, Cooperative Institute for Marine Resources Studies, Oregon State University.
2001. Co-organiser, Symposium on Research Linking Marine and Freshwater Events in Salmon Ecology and Genetics. Hatfield Marine Science Center, Newport, OR, USA.
2001. External Examiner, Ph.D dissertation, University of British Columbia, Canada.
2001. Co-editor, Proceedings of the Fourth Conference on Fish Telemetry in Europe.
2001. Invited Expert, Salmon Aquaculture Assessment Workshop, Washington, DC - organized by SeaWeb and sponsored by Rockefeller Brothers Fund.
2000. External Examiner, mid-term Ph.D. exam. University of Aarhus, Denmark.
2000. Editor, Proceedings of the Kongsvold Workshop on Releases of Salmonid Fishes in Norway. Nordic Journal of Freshwater Research.
2000. Co-organiser, Symposium on Growth Variation in Salmonids: Consequences for Life History and Implications for Management. International Congress on the Biology of Fishes, Aberdeen, Scotland.
2000. Organiser, Workshop on Releases of Anadromous Salmonids. Kongsvold, Norway.
2000. External Examiner, Ph.D. dissertation. University of Oslo, Norway.
2000. Organiser, Annual Meeting of EU Consortium. Trondheim, Norway.
1999-00. Steering Committee for the Institute Program on Effects of Ecosystem Changes on Biodiversity, Norwegian Institute for Nature Research.
1999-00. Censor, Faculty of Chemistry and Biology, Norwegian University of Science and Technology.
1998. Invited Panel Member, Regional Assessment of the Interaction between Wild and Farmed Atlantic Salmon in the Maritime Provinces, Department of Fisheries and Oceans, Canada.
1995-99. Panel Member, Norwegian Institute for Nature Research, Institute Program on Conservation of Biodiversity.
1994. Member of the International Council for the Exploration of the Sea (ICES) Study Group on Interactions of Wild, Ranched (enhanced), and Reared Salmon.

Professional Societies

1985- Society for the Study of Evolution.
1986- International Society for Behavioral Ecology.
1988-95, 02- American Fisheries Society.
1998- Fisheries Society of the British Isles.

Reviewer For

Journals: Animal Behaviour, Aquaculture, Aquaculture & Fisheries Management, Behavioral Ecology, Behavioral Ecology & Sociobiology, Canadian Journal of Fisheries & Aquatic Sciences, Canadian Journal of Zoology, Conservation Biology, Copeia, Ecology Letters, Ethology, Environmental Biology of Fishes, Evolution, Hydrobiologia, ICES Journal of Marine Science, Journal of Animal Ecology, Journal of Fish Biology, Marine Biology, Molecular Ecology, Nordic Journal of Freshwater Research, Oikos, Proceedings of the Royal Society London, Transactions of the American Fisheries Society.

Grants: Natural Environment Research Council of the United Kingdom; National Science Foundation of the USA; Pacific Sea Grant of the USA; Natural Sciences and Engineering Research Council of Canada.